[1] MCCORMICK S.Male gametophyte development[J].Plant Cell,1993,5(10):1265-1275. [2] MCCORMICK S.Control of male gametophyte development[J].Plant Cell,2004,16:S142-153. [3] 杨弘远.水稻生殖生物学[M].浙江:浙江大学出版社,2005:36-55. [4] SANDERS P M,BUI A Q,WETERINGS K,et al.Anther developmental defects in arabidopsis thaliana male-sterile mutants[J].Sexual Plant Reproduction,1999,11(6):297-322. [5] RAGHAVAN V.Anther and pollen development in rice(oryza sativa)[J].Amer J Bot,1988,75(2):183-196. [6] NONOMURA K,MIYOSHI K,EIGUCHI M,et al.The msp1 gene is necessary to restrict the number of cells entering into male and female sporogenesis and to initiate anther wall formation in rice[J].Plant Cell,2003,15(8):1728-1739. [7] ZHAO X,DE PALMA J,OANE R,et al.Ostdl1a binds to the lrr domain of rice receptor kinase msp1,and is required to limit sporocyte numbers[J].Plant J,2008,54(3):375-387. [8] JUNG K H,HAN M J,LEE D Y,et al.Wax-deficient anther1 is involved in cuticle and wax production in rice anther walls and is required for pollen development[J].Plant Cell,2006,18(11):3015-3032. [9] LI H,PINOT F,SAUVEPLANE V,et al.Cytochrome p450 family member cyp704b2 catalyzes the ω-hydroxylation of fatty acids and is required for anther cutin biosynthesis and pollen exine formation in rice[J].Plant Cell,2010,22(1):173-190. [10] HIHARA Y,HARA C,UCHIMIYA H.Isolation and characterization of two cdna clones for mrnas that are abundantly expressed in immature anthers of rice(oryza sativa.)[J].Plant Mol Biol,1996,30(6):1181-1193. [11] NGUYEN N,ZHANG X,OLASHAW N,et al.Molecular cloning and functional characterization of the transcription factor YY2[J].J Biol Chem,2004,279(24):25927-25934. [12] TSUCHIYA T,TORIYAMA K,EJIRI S,et al.Molecular characterization of rice genes specifically expressed in the anther tapetum[J].Plant Mol Biol,1994,26(6):1737-1746. [13] YOKOI S,TSUCHIYA T,TORIYAMA K,et al.Tapetum-specific expression of the osg6b promoter-β-glucuronidase gene in transgenic rice[J].Plant Cell Reports,1997,16(6):363-367. [14] DING Z J,WU X H,WANG T.The rice tapetum-specific gene RA39 encodes a type i ribosome-inactivating protein[J].Sexual Plant Reproduction,2002,15(4):205-212. [15] JUNG K H,HAN M J,LEE Y S,et al.Rice undeveloped tapetum1 is a major regulator of early tapetum development[J].Plant Cell,2005,17(10):2705-2722. [16] LUO H,LEE J Y,HU Q,et al.RTS,a rice anther-specific gene is required for male fertility and its promoter sequence directs tissue-specific gene expression in different plant species[J].Plant Mol Biol,2006,62(3):397-408. [17] LI N,ZHANG D S,LIU H S,et al.The rice tapetum degeneration retardation gene is required for tapetum degradation and anther development[J].Plant Cell,2006,18(11):2999-3014. [18] ZHANG D S,LIANG W Q,YUAN Z,et al.Tapetum degeneration retardation is critical for aliphatic metabolism and gene regulation during rice pollen development[J].Mol Plant,2008,1(4):599-610. [19] WANG A,XIA Q,XIE W,et al.Male gametophyte development in bread wheat(triticum aestivum l.):Molecular,cellular,and biochemical analyses of a sporophytic contribution to pollen wall ontogeny[J].Plant J,2002,30(6):613-623. [20] WANG A,XIA Q,XIE W,et al.The classical ubisch bodies carry a sporophytically produced structural protein(RAFTIN)that is essential for pollen development[J].PNAS,2003,100(24):14487-14492. [21] STIEGLITZ H.Role of beta-1,3-glucanase in postmeiotic microspore release[J].Dev Biol,1977,57(1):87-97. [22] CHEN R,ZHAO X,SHAO Z,et al.Rice udp-glucose pyrophosphorylase1 is essential for pollen callose deposition and its cosuppression results in a new type of thermosensitive genic male sterility[J].Plant Cell,2007,19(3):847-861. [23] YAMAGUCHI T,HAYASHI T,NAKAYAMA K,et al.Expression analysis of genes for callose synthases and rho-type small gtp-binding proteins that are related to callose synthesis in rice anther[J].Biosci Biotechnol Biochem,2006,70(3):639-645. [24] NONOMURA K,NAKANO M,FUKUDA T,et al.The novel gene homologous pairing aberration in rice meiosis1 of rice encodes a putative coiled-coil protein required for homologous chromosome pairing in meiosis[J].Plant Cell,2004,16(4):1008-1020. [25] NONOMURA K I,NAKANO M,MURATA K,et al.An insertional mutation in the rice pair2 gene,the ortholog of arabidopsis asy1,results in a defect in homologous chromosome pairing during meiosis[J].Mol Genet Genomics,2004,271(2):121-129. [26] YUAN W,LI X,CHANG Y,et al.Mutation of the rice gene pair3 results in lack of bivalent formation in meiosis[J].Plant J,2009,59(2):303-315. [27] DENG Z Y,WANG T.Osdmc1 is required for homologous pairing in oryza sativa[J].Plant Mol Biol,2007,65(1-2):31-42. [28] CHANG L,MA H,XUE H W.Functional conservation of the meiotic genes sds and rck in male meiosis in the monocot rice[J].Cell Res,2009,19(6):768-782. [29] TAO J,ZHANG L,CHONG K,et al.Osrad21-3,an orthologue of yeast rad21,is required for pollen development in oryza sativa[J].Plant J,2007,51(5):919-930. [30] ZHANG L,TAO J,WANG S,et al.The rice osrad21-4,an orthologue of yeast rec8 protein,is required for efficient meiosis[J].Plant Mol Biol,2006,60(4):533-554. [31] NONOMURA K,MOROHOSHI A,NAKANO M,et al.A germ cell specific gene of the argonaute family is essential for the progression of premeiotic mitosis and meiosis during sporogenesis in rice[J].Plant Cell,2007,19(8):2583-2594. [32] 谭何新,文铁桥,张大兵,等.水稻花粉发育的分子机理[J].植物学通报,2007,24(3):330-339. [33] 郑宏红,瞿礼嘉,刘美华,等.花药特异性表达的类查尔酮合酶基因D5与水稻花粉发育相关[J].科学通报,2000,45(11):1132-1137. [34] MORITOH S,MIKI D,AKIYAMA M,et al.Rnai-mediated silencing of osgen-l(osgen-like),a new member of the rad2/xpg nuclease family,causes male sterility by defect of microspore development in rice[J].Plant Cell Physiol,2005,46(5):699-715. [35] JIANG S Y,CAI M,RAMACHANDRAN S.The oryza sativa no pollen(osnop)gene plays a role in male gametophyte development and most likely encodes a c2-gram domain-containing protein[J].Plant Mol Biol,2005,57(6):835-853. [36] HAN M J,JUNG K H,YI G,et al.Rice immature pollen 1(rip1)is a regulator of late pollen development[J].Plant Cell Physiol,2006,47(11):1457-1472. [37] JEON J S,CHUNG Y Y,LEE S,et al.Isolation and characterization of an anther-specific gene,ra8,from rice(oryza sativa.)[J].Plant Mol Biol,1999,39(1):35-44. [38] KANEKO M,INUKAI Y,UEGUCHI-TANAKA M,et al.Loss-of-function mutations of the rice gamyb gene impair alpha-amylase expression in aleurone and flower development[J].Plant Cell,2004,16(1):33-44. [39] ZOU J T,ZHAN X Y,WU H M,et al.Characterization of a rice pollen-specific gene and its expression[J].American Journal of Botany,1994,81(5):552-561. [40] ZHU Q H,RAMM K,SHIVAKKUMAR R,et al.The anther indehiscence1 gene encoding a single myb domain protein is involved in anther development in rice[J].Plant Physiol,2004,135(3):1514-1525. [41] YANG W Q,LAI Y,LI M N,et al.A novel c2-domain phospholipid-binding protein,ospbp1,is required for pollen fertility in rice[J].Mol Plant,2008,1(5):770-785. [42] ZHAO L F,XU S B,CHAI T Y,et al.Osap2-1,an ap2-like gene from oryza sativa,is required for flower development and male fertility[J].Sex Plant Reprod,2006,19(4):197-206. [43] 江华,杨仲南,高菊芳.水稻雄性不育突变体osms121的遗传及定位分析[J].上海师范大学学报:自然科学版,2006,35(6):71-75. [44] 吴孝槐,毛爱军,王荣,等.水稻花药发育相关基因osprp1的克隆和特性分析[J].科学通报,2003,48(20):2154-5461. [45] 吴孝槐,王台.水稻花优势表达基因ra68的克隆和特性分析[J].科学通报,2004,49(13):1263-1267. |